Scientific papers using WWRG data


Clark, J.A. & Dodd, S.G. (2022) A Eurasian Oystercatcher with aberrant plumage. Wader Study 129,: 151–152. doi:10.18194/ws.00281

Cook, A.S.C.P., Burton, N.H.K., Dodd, S.G., Foster, S., Pell, R.J., Ward, R.M., Wright, L.J. & Robinson, R.A. (2021) Temperature and density influence survival in a rapidly declining migratory shorebird.  Biological Conservation 260, 109198.

Meltofte, H., Hansen, J. & Riget, F. (2021) Trends in breeding performance in wader populations at Zackenberg, high Arctic Greenland, in relation to environmental drivers 1996-2018. Polar Biology

Pell, R.J., Clark, J.A. & Robinson, R.A. (2023) Contrasting habitat use between and within Bar-tailed Godwit and Curlew wintering on the Wash. Wader Study 130, 225–233.

Schwemmer, P., Mercker, M., Haecker, K., Kruckenberg, H., Kämpfer, S., Bocher, P., Fort, J., Jiguet, F., Franks, S., Elts, J., Marja, R., Piha, M., Rousseau, P., Pederson, R., Düttmann, H., Faermann, T. & Garthe, S. (2023) Behavioral responses to offshore windfarms during migration of a declinging shorebird species revealed by GPS-telemetry. Journal of Environmental Management 118131.


Alves, J.A., Gunnarsson, T.G., Hayhow, D.B., Appleton, G.F., Potts, P.M., Sutherland, W.J. & Gill, J.A. (2013) Costs, benefits, and fitness consequences of different migratory strategies. Ecology 94, 1117.

Alves, J.A., Gunnarsson, T.G., Potts, P.M., Gélinaud, G., Sutherland, W.J. & Gill, J.A. (2012) Overtaking on migration: does longer distance migration always incur a penalty? Oikos, 121, 464470. 

Alves, J.A., Lourenço, P.M., Piersma, T., Sutherland, W.J. & Gill, J.A. (2010) Population overlap and habitat segregation in wintering Black-tailed Godwits. Bird Study 57, 381391.

Atkinson, P.W., McLean, I.M. & Clark, N.A. (2010) Impacts of shellfisheries and nutrient inputs on waterbird communities in the Wash, England. Journal of Applied Ecology 47, 191199.

Clark, N.A. & Clark, J.A. (2015) A method of making robust and effective decoys for trapping waders using dead birds.  Wader Study 122, 31-36.

Duijns, S., van Gils, J.A., Smart, J. & Piersma, T. (2015) Phenotype-limited distributions: short-billed birds move away during times that prey bury deeply. Royal Society Open Science 2:150073.

Durrel, S.E.A. le V. dit. & Atkinson, P.W. (2004) Differential distribution of Oystercatchers Haematopus ostralegus overwintering on the Wash, east England.  Bird Study 51, 76-82.

Gill, J.A. (2006) Climate change and coastal migrant birds. In Principles of Conservation Biology 3rd Edition (eds: M.J. Groom, G.K. Meffe & C.R. Carroll). Sinauer Associates, USA.

Gunnarsson, T.G., Sutherland, W.J., Alves, J.A., Appleton, G.F., Potts, P.M., & Gill, J.A. (2012)Rapid changes in phenotype distribution during range expansion in a migratory bird. Proceedings of the Royal Society of London B, 279, 411-416.


Atkinson, P.W., Clark, N.A., Bell, M.C., Dare, P.J., Clark, J.A. & Ireland, P.L. (2003) Changes in commercially fished shellfish stocks and shorebird populations on the Wash, England.  Biological Conservation 114, 127–141.

Boyd, H. & Piersma, T. (2001) Why do few Afro-Siberian Knots Calidris canutus canutus now visit Britain?  Bird Study 48, 147158.

Clark, J.A. (2002) Effects of severe weather on wintering waders. Unpublished MPhil Thesis, University of East Anglia.

Clark, J.A. (2009) Selective mortality of waders during severe weather.  Bird Study 56, 96102.

Gill, J.A., Norris, K., Potts, P.M., Gunnarsson, T.G., Atkinson, P.W. & Sutherland, W.J. (2001) The buffer effect and largescale population regulation in migratory birds.  Nature 412, 436438.

Gunnarsson, T.G., Gill, J.A., Atkinson, P.W., Gélinaud, G., Potts, P.M., Croger, R.E., Gudmundsson, G.A., Appleton, G.F. & Sutherland, W.J. (2006) Population-scale drivers of individual arrival times in migratory birds.  Journal of Animal Ecology 75, 1119–1127.

Gunnarsson, T.G., Gill, J.A., Goodacre, S.L., Gélinaud, G., Atkinson, P.W., Hewitt, G.M., Potts, P.M. & Sutherland, W.J. (2006) Sexing Black–tailed Godwits: a comparison of behavioural, molecular, biometric and field-based techniques.  Bird Study 53, 19198.

Gunnarsson, T.G., Gill, J.A., Newton, J., Potts, P.M. & Sutherland, W.J. (2005) Seasonal matching of habitat quality and fitness in migratory birds.  Proceedings of the Royal Society of London B 272, 23192323.

Gunnarsson, T.G., Gill, J.A., Potts, P.M., Atkinson, P.W., Croger, R.E. Gélinaud, G., Gardasson, A. & Sutherland, W.J. (2005) Estimating population size in Icelandic Black-tailed Godwits Limosa limosa by colour-marking.  Bird Study 52, 153158.

Gunnarsson, T.G., Gill, J.A., Sigurbjörnsson, P. & Sutherland, W.J. (2004) Arrival synchrony in migratory birds.  Nature 431, 646.

Meltofte, H. (2001) Wader population censuses in the Arctic: getting the timing right.  Arctic 54, 367376.

Robinson, R.A., Burton, N.H.K., Clark, J.A. & Rehfisch, M.M. (2007) Monitoring survival of waders in Britain. BTO Research Report 469.  BTO, Thetford. Available on the BTO website.

Serra, L., Clark, N.A. & Clark, J.A. (2006) Primary moult, body mass and migration of Grey Plovers Pluvialis squatarola in Britain.  Ibis 148, 292301.

Smart, J. & Gill, J.A. (2003) Non-intertidal habitat use by shorebirds: a reflection of inadequate intertidal resources?  Biological Conservation 111, 359369.

Verkuil, Y.I., Jukema, J., Gill, J.A., Karlionova, N., Koopman, K., Melter, J., Hooijmeijer, J.C.E.W. & Piersma, T. (2008) Nonbreeding faeders Ruffs Philomachus pugnax associate according to sex, not morphology. Bird Study, 55, 241-246.


Anderson, K.R. (1974) The biometrics, moult and recoveries of British-ringed Ruff. Wader Study Group Bulletin 11, 5-14.

Anderson, K.R. & Minton, C.D.T. (1978) Origins and movements of Oystercatchers on the Wash.  British Birds 71, 439-447.

Atkinson, P.W. (1996) The origins, moult, movements and changes in numbers of Bar-tailed Godwits Limosa lapponica on the Wash, England.  Bird Study 43, 60-72.

Bainbridge, I.P. & Minton, C.D.T. (1978) The migration and mortality of the Curlew in Britain and Ireland. Bird Study 25, 39-50.

Berry, R. (1971) Wader Studies on the Wash. Norfolk Bird and Mammal Report 1970, 176-179.

Branson, N.J.B.A. & Minton, C.D.T. (1976) Moult, measurements and migrations of the Grey Plover. Bird Study 25, 257-266.

Branson, N.J.B.A., Ponting, E.D. & Minton, C.D.T. (1978) Turnstone migration in Britain and Europe. Birdc Study 25, 181-187.

Branson, N.J.B.A., Ponting, E.D. & Minton, C.D.T. (1979) Turnstone populations on the Wash.  Bird Study 26, 47-54.

Clark, J., Clark, N. & Baillie, S. (1993) Cold comfort for waders. BTO News 186, 1.

Clark, N.A. (1978) The weights, moult and morphometrics of Spotted Redshanks in Britain. Wader Study Group Bulletin 22, 22-26.

Clark, N. (1993) Wash Oystercatchers starving. BTO News 185, 1 & 24.

Cooper, R. (1983) Painted waders. Shooting and Conservation 1983, 11.

Cooper, R.H.W. (1988) Migration strategies of shorebirds during the non-breeding season with particular reference to the Sanderling (Calidris alba).  Unpublished PhD Thesis, University of Durham.

Dick, W.J.A., Pienkowski, M.W., Waltner, M. & Minton, C.D.T. (1976) Distribution and geographical origins of Knot Calidris canutus wintering in Eutrope and Africa. Aread 64, 22-47.

Gill, J.A., Clark, J.A., Clark, N.A. & Sutherland, W.J. (1995) Sex differences in the migration, moult and wintering areas of British-ringed Ruff. Ringing & Migration 16, 159-167.

Green, G.H. (1973) Some notes on Bar-tailed Godwit ringing, biometrics and moult. Wader Study Group Bulletin 8, 4-8.

Hardy, A.R. & Minton, C.D.T. (1980) Dunlin migration in Britain and Ireland. Bird Study 27, 81-92.

Johnson, C. (1985) Patterns of seasonal weight variation in waders on the Wash.  Ringing & Migration 6, 19-32.

Johnson, C. & Minton, C.D.T. (1980) The primary moult of Dunlin at the Wash.  Ornis Scandinavica 11, 190-195.

Langston, R. (1987) The Wash – mecca for waders. Norfolk Bird and Mammal Report 1987, 105-111.

Minton, C.D.T. (1970) Wader Studies on the Wash. Norfolk Bird and Mammal Report 1969, 15-18.

Minton, C.D.T. (1977) Wader plumages in autumn and winter. British Birds 70, 521-529.

Minton, C.D.T. & Stanley, P.I. (1972) Biometric variations in the Curlew Sandpiper. Wader Study Group Bulletin 6, 7-9.

Ogilvie, M.A. (1965) Wader ringing by the Wildfowl Trust, 1959-64. The Wildfowl Trust 16th annual report, 48-54.

Pienkowski, M. (1976) Seasonal changes in bill-lengths of Knots, and a comment on bill measuring techniques for waders. Wader Study Group Bulletin 17, 12-14.

Pienkowski, M.W. & Dick, W.J.A. (1976) Some biases in cannon- and mist-netted samples of wader populations. Ringing & Migration 1, 105-107.

Pienkowski, M.W. & Minton, C.D.T. (1973) Wing length changes of Knot with age and time since moult. Bird Study 20, 63-68.

Pienkowski, M.W., Lloyd, C.S. & Minton, C.D.T. (1979) Seasonal and migrational weight changes in Dunlins. Bird Study 26, 134-148.

Rehfisch, M.M., Clark, N.A., Langston, R.H.W. & Greenwood, J.J.D. (1996) A guide to the provision of refuges for waders: an analysis of 30 years of ringing data from the Wash, England. Journal of Applied Ecology 33, 673-687.

Stanley, P.I. & Minton, C.D.T. (1972) The unprecedented westward migration of Curlew Sandpipers in autumn 1969. British Birds 65, 365-380.

Historical documents

Branson, N.J.B.A. (1978) The work of the Wash Wader Ringing Group. Paper contributing to the Wash Symposium organised by the Lincolnshire and South Humberside Trust for Nature Conservation.

Ireland, P. & Clark, N. (1989) A thirtieth birthday on the Wash. BTO News 164, 7.

Minton, C.D.T. (1971) Wader ageing guide.

Minton, C.D.T. (1976?) The waders of the Wash – Ringing and biometric studies. Wash feasibility study. Ecological Report. Scientific Study G.

Minton, C.D.T. & Boyd, H. (??) Age, wing moult and measurements of waders on the Wash in late summer. Two drafts of a paper?

NERC (1976) The Wash feasibility study – a report on the ecological studies. NERC Publication series C.

Pienkowski, M.W. (1983) Identification of relative importance of sites by studies of movement and population turnover. Workshop contribution. Also reports from the Movements Project.

Pienkowski, M.W. & Minton, C.D.T. (ca. 1973) The use of wing length in separating bird populations. Draft of Knot wing lengths paper.

WWRG (1969) Cannon nets – construction, operation and application of equipment used in the United Kingdom by the Wash Wader Ringing Group. Unpublished Booklet.